Interesting points!
If you look at the protein data alone, there are no recovered variants that support these samples being very divergent from modern humans, e.g. 'H.erectus like'. For comparison, the H.antecessor enamel had a variant, placing it as an outgroup to nean, deni, and present day humans
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High-throughput paleoproteomics method using MALDI-CASI-FTICR mass spectrometry to estimate biological sex from tooth enamel.
Great success on 130 individuals from medieval Great Moravia.
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The two variants that are recovered are also found on the same protein!
If they were introgressed from homo erectus, they likely would have been linked together due to their physical proximity on the genome, and Denisovans would have inherited both.
While we suspect incomplete lineage sorting as the main culprit for this result, an older admixture between Neanderthal and humans, a 'super-archaic' admixture into Denisovans or other scenarios could be at play.