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Abril, S. & Gómez, C. (2009). Ascertaining key factors behind the coexistence of the native ant species Plagiolepis pygmaea with the invasive Argentine ant Linepithema humile (Hymenoptera: Formicidae). Sociobiology, 53: 559-568.
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Non-native Ants of the Iberian Peninsula: a 2025 Update | Sociobiology