Interesting – this is what you need for synaptic plasticity to solve "within-neuron" credit assignment. Backpropagating APs, elevating Ca2+ above the plasticity threshold, should preferentially enter dendrites which recently activated.
Is this the first evidence of this?
Ari Benjamin
But then where do the branch-specific variations in voltage come from? We found that in apical dendrites, small subthreshold depolarizations *preceding* a complex spike partially predict voltage (and Ca2+) amplitude *during* the event. This suggests conjunctive amplification of inputs and spikes.