Our paper unpacking "fitness" is out doi.org/10.1093/gene.... How to define fitness depends on what question you ask. Selection acts on organismal vital rates, longer-term outcomes make sense for genetic lineages, models link them. "Invasion fitness" = speed, but sometimes probability matters more.
Abstract. Evolutionary “fitness” is operationalized in many different ways in models. Its role is to quantify that which is favored by natural selection. G
Our latest preprint doi.org/10.32942/X2V... explains why fitness can be defined in so many ways, and which version(s) you should use when. Fitness quantifies what natural selection favors. 1/